Internal Pedometer of Cataglyphis

0n June 29th 2006, The Economist published an article entitled 'A stilted story'. This article discusses how Dr. Matthias Wittlinger manipulated ant appendages 'to investigate a century-old hypothesis that desert ants have internal pedometers'. Here is an excerpt:

If there were a Royal Society for the Prevention of Cruelty to Ants, Matthias Wittlinger of the University of Ulm, in Germany, would probably be top of its hate list. The reason is that Dr Wittlinger and his colleagues have, as they report in this week's Science, been chopping the feet off ants. And not only that. They have been making other ants walk around on stilts.

Saharan desert ants of the genus Cataglyphis have to travel long distances to discover food in their impoverished, sandy environment. How they find their way home once they have done so is a mystery. Ants in more temperate climates often lay down chemical trails, but Cataglyphis, apparently, does not. Like honeybees and ancient mariners, they can navigate by the sun, so they know the general direction in which to travel. But, also like ancient mariners (who knew their latitude, but not their longitude), such solar reckoning cannot tell them when to stop.

Dr Wittlinger, therefore, decided to investigate a century-old hypothesis that desert ants have internal pedometers—in other words, they count their steps out, and they count them back. When one total matches the other, they are home.